Zeora symmicta (Ach.) L. M. Weber & Ivanovich-Hichins

in Ivanovich-Hichins & al., Lichenologist, 57: 300, 2025. Basionym: Lecanora varia var. symmicta Ach. - Lichenogr. Univ.: 379, 1810.
Synonyms: Biatora maculiformis (Hoffm.) Beltr.; Biatora symmicta (Ach.) A. Massal.; Lecanora symmicta (Ach.) Ach.; Lecanora symmicta var. symmictera (Nyl.) Zahlbr.; Lecanora symmictera Nyl.; Lecanora trabalis (Ach.) Nyl.; Lecanora varia var. maculiformis (Hoffm.) Rabenh.; Lecidea symmicta (Ach.) Ach.; Zeora maculiformis (Hoffm.) Trevis.
Description: Thallus crustose, continuous, rimose or warted-areolate, sometimes almost subleprose, yellowish green to greenish grey, sometimes inapparent, rarely delimited by a white prothallus. Apothecia biatorine, round to irregular in outline, sometimes confluent and/or tuberculate, sessile, 0.4-0.7(-1) mm across, with a yellowish green to ochre-coloured, rarely darkened, weakly to strongly convex disc and thin, soon excluded proper margin, without a thalline margin. Proper exciple colourless within, the outer part with yellow-brown granules soluble in K, of strongly gelatinized, branched and anastomosing, radiating hyphae with narrowly cylindrical lumina; epithecium greenish ochre to orange-brown, with brownish yellow, coarse granules soluble in K, 5-15 µm high; hymenium colourless to pale yellow-brown, 35-55 µm high; paraphyses sparingly branched and anastomosing, 1.5-2.5 µm thick at mid-level, the apical cells not much swollen, up to 3.5 µm wide; hypothecium colourless. Asci 8-spored, clavate, very thin-walled, with a K/I+ blue, tall tholus penetrated by a faintly amyloid apical cushion, the wall K/I-, surrounded by a blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, narrowly ellipsoid, (7-)9-12(-16) x 3-4.5(-6) µm. Conidia thread-like, 18-25 x 0.5-1 µm. Photobiont chlorococcoid. Spot tests: thallus K-, C- or C+ orange, KC- or KC+ weakly yellow, P-, UV+ dull orange, K/UV(wet)+ bright yellow-green. Chemistry: usnic acid, zeorin, arthothelin, thiophanic acid, variable amounts of xanthones.
Growth form: Crustose
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: rare
Montane belt: rather rare
Dry submediterranean belt: rather common
Humid submediterranean belt: rather common
Padanian area: extremely rare

pH of the substrata:

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Rarity

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Predictive map

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php . Courtesy: A. Stridvall


Harry Taylor; Owner: Natural History Museum London


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003


Felix Schumm – CC BY-SA 4.0
[11045], Portugal, Azoren, Sao Miguel, Südufer des Lagoa das Furnas, 270 m, 37°45.022' N, 25°19.807' W . Leg. Schumm 03.06.2003, det. Schumm 2003
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start, 4: Position von Norstictinsäure, 7: Position von Atranorin us: Usninsäure, z: Zeorin


Photo by Maria Zardini, Museo di Storia Naturale di Venezia CC BY-SA 4.0
ad cort. Abietum in nemoribus Septem Communium.
Trevisan Lichenotheca Veneta 65 as Zeora maculiformis


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.


P.L. Nimis CC BY-SA 04
TSB 14348


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Ragnitz
04.2015


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Ragnitz
04.2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, sur Pinus - La chapelle Genest, bois de Celles - Haute-Loire
19/07/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, sur Pinus - La chapelle Genest, bois de Celles - Haute-Loire
19/07/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, sur Pinus - La chapelle Genest, bois de Celles - Haute-Loire
19/07/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, sur Pinus - La chapelle Genest, bois de Celles - Haute-Loire
19/07/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, Région d'Ajaccio, sur piquet en châtaignier - Corse
16/10/2018


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, Région d'Ajaccio, sur piquet en châtaignier - Corse
16/10/2018


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_3/Lecanora_symmicta.htm
France, Région d'Ajaccio, sur piquet en châtaignier - Corse
16/10/2018


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Lesteven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Sillon de Talbert


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Sillon de Talbert


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Sillon de Talbert


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Sillon de Talbert


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Kerdreux


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
France, Kerdreux


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=458&lang=en
Belgium, Couvin


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria