Melanelia stygia (L.) Essl.
Mycotaxon, 7: 47, 1978. Basionym: Lichen stygius L. - Sp. Pl., 2: 1143, 1753.
Synonyms: Cetraria stygia (L.) Schaer.; Cornicularia stygia (L.) Nyl.; Imbricaria stygia (L.) DC.; Lichen fahlunensis L.; Parmelia fahlunensis var. stygia (L.) Schaer.; Parmelia reagens (Servít) Gyeln.; Parmelia stygia (L.) Ach.
Description: Thallus foliose, heteromerous, dorsiventral, adnate to ascending, forming 3-6(-10) cm wide rosettes. Lobes 1-5 mm long, (0.5-)1-2(-3) mm wide, convex (flattened in upper part), contiguous to imbricate, with thickened and usually down-turned margins. Upper surface dark brown to black (greenish brown in shade-forms), more or less glossy, with rounded to elongate, whitish, mostly marginal pseudocyphellae which are often surrounded by a raised, often darker margin; lower surface dark brown to black, matt, with scattered, mostly simple, black rhizines. Upper cortex paraplectenchymatous, with a non-pored epicortex, the cell walls containing isolichenan; medulla white; lower cortex paraplectenchymatous. Apothecia lecanorine, mostly submarginal, (2-)4-6(-8) mm across, with a dark disc and a usually knobby, pseudocyphellate thalline margin. Epithecium brownish; hymenium and hypothecium colourless. Asci 8-spored, broadly clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, broadly ellipsoid, 7-12 x 4-6 µm. Pycnidia frequent, usually numerous, black, laminal, globose, mostly immersed, often with a gaping ostiole. Conidia dumbbell-shaped (thickened at both ends), 3.5-5.5 x c. 1 µm. Photobiont chlorococcoid. Spot tests: upper cortex K-, C-, KC-, P-, N-; medulla K+ yellow turning dirty red-brown (rarely K-), C-, KC-, P+ orange-red (rarely P-), I-. Chemistry: several chemotypes, with 1) stictic and norstictic acids (major), 2) fumarprotocetraric and protocetraric acids (major), 3) no lichen substances. Caperatic, cryptostictic, menegazziaic and constictic acids have been also reported as minor substances.
Growth form: Foliose, narrow lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: very common
Subalpine belt: rather common
Montane belt: very rare
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Mark Rahill - CC BY-SA 4.0
USA
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35210)
2002/07/09
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35210)
2002/07/09
pseudocyphellae
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (6845)
2001/12/05
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
c: caperatic acid
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
c: caperatic acid
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
f: fumarprotocetraric acid, p: protocetraric acid, a & b unknowns
P.L. Nimis - CC BY-SA 4.0
TSB 41998
P.L. Nimis - CC BY-SA 4.0
TSB 41999
P.L. Nimis - CC BY-SA 4.0
TSB 41999
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Pier Luigi Nimis - CC BY-SA 4.0
TSB 43972
Pier Luigi Nimis - CC BY-SA 4.0
TSB 43972
Pier Luigi Nimis - CC BY-SA 4.0
TSB 43972
Pier Luigi Nimis - CC BY-SA 4.0
TSB 44004
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 4/8/2011 - Mont Cenis - 2081 m - Savoie
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 4/8/2011 - Mont Cenis - 2081 m - Savoie
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Käsplatte
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Käsplatte
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Plöckenstein
Wolfgang von Brackel - CC BY-4.0
Czech Republic, Sumava Frymburk
Wolfgang von Brackel - CC BY-4.0
Czech Republic, Sumava Frymburk
Triin Aimla; Owner: University of Tartu
Growth form: Foliose, narrow lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: very common
Subalpine belt: rather common
Montane belt: very rare
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent
pH of the substrata:
| 1 | 2 | 3 | 4 | 5 |
Solar irradiation:
| 1 | 2 | 3 | 4 | 5 |
Aridity:
| 1 | 2 | 3 | 4 | 5 |
Eutrophication:
| 1 | 2 | 3 | 4 | 5 |
Poleotolerance:
| 0 | 1 | 2 | 3 |
Altitudinal distribution:
| 1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Mark Rahill - CC BY-SA 4.0
USA
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35210)
2002/07/09
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35210)
2002/07/09
pseudocyphellae
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (6845)
2001/12/05
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
Felix Schumm - CC BY-SA 4.0
[20334], U.R.S.S. Cucasus Magnus, ad latera occidentalia montis Elbrus, loco Krugozor dicto, 3000-3100 m, ad saxa vulcanica. Leg. A. Vezda, 21.06.1980. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1539
c: caperatic acid
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
c: caperatic acid
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
Felix Schumm - CC BY-SA 4.0
[20335], Bohemoslovacia, Slovacia centr., montes Štiavnické vrchy, in Monte Kamenná loco dicto Kamenné supra pagum Vyhne, 370 m, ad saxa eruptiva. Leg. I. Pišút et M. Svetlíková, 7.08.1987. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 2194
f: fumarprotocetraric acid, p: protocetraric acid, a & b unknowns
P.L. Nimis - CC BY-SA 4.0
TSB 41998
P.L. Nimis - CC BY-SA 4.0
TSB 41999
P.L. Nimis - CC BY-SA 4.0
TSB 41999
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Pier Luigi Nimis - CC BY-SA 4.0
TSB 43972
Pier Luigi Nimis - CC BY-SA 4.0
TSB 43972
Pier Luigi Nimis - CC BY-SA 4.0
TSB 43972
Pier Luigi Nimis - CC BY-SA 4.0
TSB 44004
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 4/8/2011 - Mont Cenis - 2081 m - Savoie
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 4/8/2011 - Mont Cenis - 2081 m - Savoie
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanelia_stygia.htm
France, 6/10/2011 - Restonica - Corse
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Käsplatte
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Käsplatte
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Plöckenstein
Wolfgang von Brackel - CC BY-4.0
Czech Republic, Sumava Frymburk
Wolfgang von Brackel - CC BY-4.0
Czech Republic, Sumava Frymburk
