Lepra amara (Ach.) Hafellner

in Hafellner & Türk, Stapfia, 104: 171, 2016. Basionym: Variolaria amara Ach. - K. Vetensk.-Akad. Nya Handl., 30, 3: 163, 1809.
Synonyms: Marfloraea amara (Ach.) S. Y. Kondr., L. Lőkös & J.-S. Hur; Pertusaria amara (Ach.) Nyl. var. amara; Pertusaria amara f. isidiata Harm.; Pertusaria communis var. amara (Ach.) Rabenh.; Pertusaria faginea auct. ital. p.p. non (L.) Leight.
Description: Thallus crustose, episubstratic, continuous to fissured, whitish, pale grey or pale greenish grey, often weakly zoned at margin, delimited by a white prothallus, sorediate, tasting extremely bitter. Central parts of thallus with numerous, circular, 0.5-1.5(-2) mm wide, white, usually convex to hemispherical soralia which long remain discrete and are sometimes arranged in concentric lines, rarely becoming confluent in central parts. In some forms the thallus is denseluy covered in warts that break down apically into small, elevated, punctiform soralia, never forming discrete round soralia. Apothecia extremely rare, immersed in soralia-like warts and not visible without sectioning the warts, the disc densely white-pruinose. Hymenium colourless, the hymenial gel not or only weakly amyloid; paraphyses lax, branched and richly anastomosing; hypothecium colourless. Asci 1-spored, broadly cylindrical, the apex without a distinct ocular chamber, the outer sheath K/I+ blue, otherwise K/I–, with an inner extensible layer, Pertusaria-type. Ascospores 1-celled, hyaline, ellipsoid, 130-150 x 40-50 µm. Photobiont chlorococcoid. Spot tests: cortex K- or K+ yellow turning orange-red, C-, KC-, P- or P+ yellow to red; soralia and medulla K- or K+ yellow turning orange-red, C-, KC+ fleeting violet, P- or P+ yellow to red, UV-. Chemistry: picrolichenic acid; a chemotype with additional protocetraric and conprotocetraric acids is also widespread.
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: rather rare
Montane belt: very common
Dry submediterranean belt: common
Humid submediterranean belt: very common
Padanian area: extremely rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
rare
rather rare
rather common
common
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extremely common
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

Nere Sorrentino
Spain, Galicia, Pontevedra


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34864)
2002/02/16


Andres Saag; Owner: University of Tartu


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36782)
2008.02.28


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36782)
2008.02.28


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36782)
2008.02.28


Antonio Valero; Owner: Antonio Valero
Spain
2010


Antonio Valero; Owner: Antonio Valero
Spain
2010


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Felix Schumm - CC BY-SA 4.0
[1320], Germany, Baden-Württemberg, Kreis Göppingen, Marbachtal bei Zell-Börtlingen, an Quercus, ca. 350 m, TK 7223. Leg. et det. F. Schumm 01.10.1967.


Felix Schumm - CC BY-SA 4.0
[1320], Germany, Baden-Württemberg, Kreis Göppingen, Marbachtal bei Zell-Börtlingen, an Quercus, ca. 350 m, TK 7223. Leg. et det. F. Schumm 01.10.1967.


Felix Schumm - CC BY-SA 4.0
[1320], Germany, Baden-Württemberg, Kreis Göppingen, Marbachtal bei Zell-Börtlingen, an Quercus, ca. 350 m, TK 7223. Leg. et det. F. Schumm 01.10.1967.
1: picrolichenic acid, 2: norstictic acid, 3: protocetraric acid, conf. J.A. Elix


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Pier Luigi Nimis - CC BY-SA 4.0
TSB 37587


P.L. Nimis CC BY-SA 4.0
TSB 37587


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_4/Lepra_amara.htm
France, sur écorce - session AFL 2005 en Lozère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_4/Lepra_amara.htm
France, sur écorce - session AFL 2005 en Lozère


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=977&lang=en
France, Daoulas


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=977&lang=en
France, Daoulas


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=977&lang=en
France, Daoulas


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=977&lang=en
France, Daoulas


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=977&lang=en
France, Menez-Meur


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=977&lang=en
France, Lesteven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1268&lang=en
France, Le Juc'h
on an enamelled iron road sign


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1268&lang=en
France, Le Juc'h
on an enamelled iron road sign


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1268&lang=en
France, Le Juc'h
on an enamelled iron road sign


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1268&lang=en
France, Le Juc'h
on an enamelled iron road sign


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1268&lang=en
France, Le Juc'h
on an enamelled iron road sign


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.


Felix Schumm - CC BY-SA 4.0
[13626], Germany, Baden-Württemberg, Kreis Göppingen, südwestlich von Adelberg nahe dem Herrenbachstausee, Buhwiesenweg, an Laubbaum im Mischwald, ca. 48°44'N, 9°33'E, 412 m, TK: 7223. Leg. et det. F. Schumm, 08.02.2008. - Sorale K-, C-, KC+ violett, P+ rostrot, UVL+ düster grünlichgelb, UVK


Felix Schumm - CC BY-SA 4.0
[13626], Germany, Baden-Württemberg, Kreis Göppingen, südwestlich von Adelberg nahe dem Herrenbachstausee, Buhwiesenweg, an Laubbaum im Mischwald, ca. 48°44'N, 9°33'E, 412 m, TK: 7223. Leg. et det. F. Schumm, 08.02.2008. - Sorale K-, C-, KC+ violett, P+ rostrot, UVL+ düster grünlichgelb, UVK


Felix Schumm - CC BY-SA 4.0
[13626], Germany, Baden-Württemberg, Kreis Göppingen, südwestlich von Adelberg nahe dem Herrenbachstausee, Buhwiesenweg, an Laubbaum im Mischwald, ca. 48°44'N, 9°33'E, 412 m, TK: 7223. Leg. et det. F. Schumm, 08.02.2008. - Sorale K-, C-, KC+ violett, P+ rostrot, UVL+ düster grünlichgelb, UVK


Felix Schumm - CC BY-SA 4.0
[13626], Germany, Baden-Württemberg, Kreis Göppingen, südwestlich von Adelberg nahe dem Herrenbachstausee, Buhwiesenweg, an Laubbaum im Mischwald, ca. 48°44'N, 9°33'E, 412 m, TK: 7223. Leg. et det. F. Schumm, 08.02.2008. - Sorale K-, C-, KC+ violett, P+ rostrot, UVL+ düster grünlichgelb, UVK


Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain


Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain


Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Schweinfurt