Phlyctis agelaea (Ach.) Flot.

Bot. Zeit., 8: 574, 1850. Basionym: Lichen agelaeus Ach. - Lichenogr. Suec. Prodr.: 30, 1799.
Synonyms: Phlyctis italica Garov.; Phlyctis spilomatica (Schaer.) A. Massal. ?
Description: Thallus crustose, ecorticate, thinly episubstratic, continuous or more or less cracked or verrucose, white to grey-white, forming up to 5(-12) cm wide patches delimited by a white prothalline line. Apothecia deeply immersed, clustered by 2-4 at the center of flattened, irregular, powdery or granular, 0.2-0.5(-1) mm wide thalline warts which are usually mistaken for soralia; disc concave to flat, brown-grey to reddish brown, but covered by the granular warts; thalline margin white-granular. Proper exciple thin, grey, I+ pale blue, usually obscured by the thalline exciple; epithecium colourless to brownish, strongly granular; hymenium colourless, K/I+ blue; paraphyses simple or sparingly branched in upper part, not capitate, c. 1.5 μm wide at base, c. 2.5 μm at apex; hypothecium pale brown. Asci 2(-4)-spored, broadly clavate, thin-walled, not strongly thickened at apex, the outer wall K/I+ blue, the tholus I+ weakly blue. Ascospores strongly muriform, hyaline to finally yellow-brown, broadly ellipsoid, clearly apiculate, (35-)45-80(-85) x (12-)14-30(-35) µm, with a thin, I+ blue wall and a very thin perispore. Pycnidia immersed, black. Conidia simple, ellipsoid, straight to rarely curved, 6-8 x 3-4 µm. Photobiont chlorococcoid. Spot tests: K+ yellow then slowly blood-red (needle-like crystals), C-, KC+ reddish, P+ yellow then orange-red, UV-. Chemistry: norstictic acid.
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: extremely rare
Dry submediterranean belt: rare
Humid submediterranean belt: rather common
Padanian area: absent

pH of the substrata:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

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Predictive map

Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11012)
2001/11/28


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11012)
2001/11/28


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.


Manuel Gil
Spain, Canary Islands, Tenerife


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 2012, session Normandie, gorges du Meillon, Le Vaudobin


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Landevennec


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, St. Julien


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon


Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento


Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento


Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento


Felix Schumm - CC BY-SA 4.0
[VZR448], Italia. Insula Sardinia, Cagliari, reservatum naturae dictum Monte Arcosu, prope domum forestalem Casa Perdu Molis, 500 m. Ad corticem Quercus Ilicis. Leg. & det. A. Vezda, 29.8.1989. Ex A. Vezda: Lichenes Rariores Exsiccati Nr. 448.


Felix Schumm - CC BY-SA 4.0
[VZR448], Italia. Insula Sardinia, Cagliari, reservatum naturae dictum Monte Arcosu, prope domum forestalem Casa Perdu Molis, 500 m. Ad corticem Quercus Ilicis. Leg. & det. A. Vezda, 29.8.1989. Ex A. Vezda: Lichenes Rariores Exsiccati Nr. 448.


Felix Schumm - CC BY 4.0
[VZ1112], Romania. Distr. Hunedoara, Montes Retezat, in valle Lapu ul Mare, supra Gura Apei, 1050 m. Ad corticem Aceris pseudoplatani. Leg. A. Vezda, 5.6.1972. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1112.


Felix Schumm - CC BY 4.0
[VZ1112], Romania. Distr. Hunedoara, Montes Retezat, in valle Lapu ul Mare, supra Gura Apei, 1050 m. Ad corticem Aceris pseudoplatani. Leg. A. Vezda, 5.6.1972. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1112.


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.