Collemopsis schaereri (A. Massal.) Cromb.

J. Bot., London, 12: 332, 1874.. Basionym: Pannaria schaereri A. Massal. - Ric. Auton. Lich. Crost.: 114, 1852.
Synonyms: Collema subbadium Nyl.; Collemopsis caesia Nyl.; Psorotichia caesia (Nyl.) Forssell; Psorotichia schaereri (A. Massal.) Arnold; Pyrenopsis schaereri (A. Massal.) Nyl.; Trachyderma schaereri (A. Massal.) Trevis.
Description: Thallus crustose, areolate, homoiomerous, paraplectenchymatous throughout, the hyphae small-celled, 4-9 µm thick, forming a dense network, attached by rhizohyphae. Areoles irregularly shaped, 0.2-1 mm wide, c. 0.2-0.25 mm thick, contiguous, dull dark green to black, rarely grey-pruinose, densely covered by isidia-like granules. Apothecia at first immersed then sessile, 1 or rarely 2 per areole, 0.25-0.7 mm across, zeorine, with an initially punctiform, later expanded, reddish brown, somehow glossy disc surrounded by a thin, often pale yellowish proper margin and a thick, granulose thalline margin. Proper exciple 7.5-10(-15) µm wide, cupuliform, prosoplectenchymatous, thicker in upper part; epithecium orange-brown, 10-15 µm high; hymenium hyaline, (75-)90-110(-150) µm high, I+ blue; paraphyses branched and anastomosing, with cylindrical cells, 2-3 µm thick, only slightly constricted at septa; hypothecium colourless, up to 40 µm high. Asci 8-spored, cylindrical-clavate, with a rather thick, not amyloid wall, without apical amyloid thickenings, 65-80 x 14-23 µm. Ascospores 1-celled, hyaline, ellipsoid, (9-)15-20(-25) x 6-10 µm. Pycnidia immersed to slightly projecting, globose to broadly pyriform, 50-90 µm across, with simple conidiophores. Conidia broadly to narrowly ellipsoid, (2-)3-4 x 1-2 µm, produced terminally. Photobiont cyanobacterial, chroococcoid, of single cells measuring 5-8(-10) x 5-7 µm, surrounded by a gelatinous sheath which is yellowish brown near the thallus surface. Spot tests: all negative. Chemistry: without lichen substances.
Growth form: Crustose
Substrata: rocks
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual
On otherwise dry surfaces with short periods of water seepage after rain

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: extremely rare
Montane belt: very rare
Dry submediterranean belt: rare
Humid submediterranean belt: rather rare
Padanian area: absent

pH of the substrata:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

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Occurrence data
Predictive map

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (32362)
2001/12/10


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Psorotichia_schaereri.htm
France, 30/03/2018 - Lamothe-Landerron, alt. 32 m - Gironde - (33) - au pied des murs de l'église


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Felix Schumm - CC BY-SA 4.0
[ABL51739], Taiwan,Taichung County, 15 km E of Taichung, citrus garden, on sandstone, 24°9’20’’ N, 120°49’45’’ E, 450 m. Leg. A. Aptroot (no 51739), 7.10.2001, det. A. Aptroot, 2002.


Felix Schumm - CC BY-SA 4.0
[ABL51739], Taiwan,Taichung County, 15 km E of Taichung, citrus garden, on sandstone, 24°9’20’’ N, 120°49’45’’ E, 450 m. Leg. A. Aptroot (no 51739), 7.10.2001, det. A. Aptroot, 2002.


Felix Schumm - CC BY-SA 4.0
[16648], Germany, Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392° N, 7.961653° E, 275 m, an schattiger Mauer. Leg. Schumm, 02.09.2010, det. M.Schultz. Sporen hyalin, 1-zellig, 14.6-18.7 × 6.6-7.5 μm.


Source: Prieto M., Wedin M., Schultz M. 2024. Phylogeny, evolution and a re-classification of the Lichinomycetes. Studies in Mycology, 109: 595-655. - CC BY-NC-ND
Thallus black, areoles with adnate apothecia (Schultz 08926b) Scale bar: 1 mm


Source: Prieto M., Wedin M., Schultz M. 2024. Phylogeny, evolution and a re-classification of the Lichinomycetes. Studies in Mycology, 109: 595-655. - CC BY-NC-ND
Grey pruinose morphotype, apothecia with reddish brown, open discs (Teuber 1527). Scale bar: 1 mm


Source: Prieto M., Wedin M., Schultz M. 2024. Phylogeny, evolution and a re-classification of the Lichinomycetes. Studies in Mycology, 109: 595-655. - CC BY-NC-ND
Apothecium with zeorine margin, subhymenium inversely cone shaped in LPCB (Massalongo Lich. Ital. 338, UPS lectotype). Scale bar: 25 micron


Source: Prieto M., Wedin M., Schultz M. 2024. Phylogeny, evolution and a re-classification of the Lichinomycetes. Studies in Mycology, 109: 595-655. - CC BY-NC-ND
D. Zeorine apothecial margin (Schultz 03438). E. Paraplectenchymatous thallus anatomy in LPCB (Schultz 03440b). Scale bars: C = 25 µm; D, E = 10 µm.


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, in op. Tregnago ad saxa irrigua 1855
as Psorotichia schaereri