Pycnothelia papillaria Dufour

Ann. Gén. Sci. Phys., 8: 46, 1821.
Synonyms: Biatora epimarta (Nyl.) Walt. Watson; Cenomyce papillaria Ach. nom. illegit.; Cladonia papillaria Hoffm nom. illegit.; Cladonia trapezuntica J. Steiner; Lecidea epimarta Nyl.
Description: Primary thallus crustose, persistent (rarely disappearing in exceptionally well-developed specimens), of ecorticate, greenish grey or whtish granules grouped into 20-40 mm wide rosettes. Podetia wart-like to short-cylindrical, cream-coloured, grey to brownish, hollow inside, 2-5(-10) mm tall, constricted at base, smooth, esquamulose, with rounded, darkened apices, mostly simple, but sometimes sparingly branched in upper part, with frequent, dark pycnidial ostioles at the tips, the internal surface cartilaginous and striate, <0.5 mm thick. Apothecia very rare, terminal, brown, convex, emarginate, 0.3-1 mm across. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores simple or 1(-3)-septate, colourless, fusiform, thin-walled, (7-)9-15 x 2-4(-5) µm. Pycnidia dark, more or less immersed, with a colourless slime. Conidia filiform, curved, 8–15 × c. 0.5 µm. Photobiont chlorococcoid. Spot tests: thallus K+ yellow, C-, KC-, P-; medulla UV+ blue-white. Chemistry: atranorin, and variable amounts of chloratranorin and lichesterinic, protolichesterinic and squamatic acids.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: rather rare
Montane belt: very rare
Dry submediterranean belt: extremely rare
Humid submediterranean belt: extremely rare
Padanian area: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
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common
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11504)
2001/11/29


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Camaret


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36806)
2008.02.25


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019


Mark Rahill - CC BY-SA 4.0
USA


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre


P.L. Nimis CC BY-SA 4.0
TSB 38871


P.L. Nimis CC BY-SA 4.0
TSB 42077


P.L. Nimis CC BY-SA 4.0
TSB 35215


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Piemonte, Biella, Val Sessera, 450 m
12/2018
very well-developed specimens, with almost no primary thallus


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Piemonte, Biella, Val Sessera, 450 m
12/2018
very well-developed specimens, with almost no primary thallus


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Kerloc'h


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Kerloc'h


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Telgruc


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Camaret


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria Tennenlohe


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Arber


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Grafenwöhr


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Grafenwöhr


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Leinburg


Wolfgang von Brackel - CC BY-4.0
Italy, Liguria, Passo della Forcella


Wolfgang von Brackel - CC BY-4.0
Italy, Liguria, Passo della Forcella


Triin Aimla; Owner: University of Tartu


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.