Pycnothelia papillaria Dufour
Ann. Gén. Sci. Phys., 8: 46, 1821.
Synonyms: Biatora epimarta (Nyl.) Walt. Watson; Cenomyce papillaria Ach. nom. illegit.; Cladonia papillaria Hoffm nom. illegit.; Cladonia trapezuntica J. Steiner; Lecidea epimarta Nyl.
Description: Primary thallus crustose, persistent (rarely disappearing in exceptionally well-developed specimens), of ecorticate, greenish grey or whtish granules grouped into 20-40 mm wide rosettes. Podetia wart-like to short-cylindrical, cream-coloured, grey to brownish, hollow inside, 2-5(-10) mm tall, constricted at base, smooth, esquamulose, with rounded, darkened apices, mostly simple, but sometimes sparingly branched in upper part, with frequent, dark pycnidial ostioles at the tips, the internal surface cartilaginous and striate, <0.5 mm thick. Apothecia very rare, terminal, brown, convex, emarginate, 0.3-1 mm across. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores simple or 1(-3)-septate, colourless, fusiform, thin-walled, (7-)9-15 x 2-4(-5) µm. Pycnidia dark, more or less immersed, with a colourless slime. Conidia filiform, curved, 8–15 × c. 0.5 µm. Photobiont chlorococcoid. Spot tests: thallus K+ yellow, C-, KC-, P-; medulla UV+ blue-white. Chemistry: atranorin, and variable amounts of chloratranorin and lichesterinic, protolichesterinic and squamatic acids.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: rare
Subalpine belt: rather rare
Montane belt: very rare
Dry submediterranean belt: extremely rare
Humid submediterranean belt: extremely rare
Padanian area: absent
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
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Occurrence data
Predictive map
Current prediction (1981-2010)
Future prediction (2071-2100) SSP 1-2.6
Future prediction (2071-2100) SSP 5-8.5Predictive maps according to Francesconi et al. 2025
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11504)
2001/11/29

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Camaret

P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36806)
2008.02.25

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Piemonte, Biella, Val Sessera, 450 m
12/2018
very well-developed specimens, with almost no primary thallus

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Piemonte, Biella, Val Sessera, 450 m
12/2018
very well-developed specimens, with almost no primary thallus

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Kerloc'h

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Kerloc'h

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Telgruc

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Camaret

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: rare
Subalpine belt: rather rare
Montane belt: very rare
Dry submediterranean belt: extremely rare
Humid submediterranean belt: extremely rare
Padanian area: absent
pH of the substrata:
1 | 2 | 3 | 4 | 5 |
Solar irradiation:
1 | 2 | 3 | 4 | 5 |
Aridity:
1 | 2 | 3 | 4 | 5 |
Eutrophication:
1 | 2 | 3 | 4 | 5 |
Poleotolerance:
0 | 1 | 2 | 3 |
Altitudinal distribution:
1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map



Predictive maps according to Francesconi et al. 2025

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11504)
2001/11/29

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Camaret

P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36806)
2008.02.25

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Piemonte, Biella, Val Sessera, 450 m
12/2018
very well-developed specimens, with almost no primary thallus

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Piemonte, Biella, Val Sessera, 450 m
12/2018
very well-developed specimens, with almost no primary thallus

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Cap de la Chèvre

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Kerloc'h

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Kerloc'h

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Telgruc

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=251&lang=en
France, Camaret

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Santa María del Puerto (Somiedo-Asturias), 16-VII-2024, sobre suelo ácido, leg. & det. M. González, MGG-126.
