Ricasolia amplissima (Scop.) De Not. - chloromorph

Giorn. Bot. Ital., 2: 179, 1846. Basionym: Lichen amplissimus Scop. - Fl. Carniol., 2 ed.: 386, 1772.
Synonyms: Lobaria amplissima (Scop.) Forssell; Lobaria glomulifera (Lightf.) Hoffm.; Lobaria laciniata (Huds.) Vain.; Parmelia amplissima (Scop.) Schaer.; Parmelia glomulifera (Lightf.) Ach.; Ricasolia glomulifera (Lightf.) Nyl.; Sticta amplissima (Scop.) Rabenh.; Sticta glomulifera (Lightf.) Delise
Description: Thallus foliose, heteromerous, dorsiventral, broad-lobed, more or less coriaceous, 0.5-0.7 mm thick, usually forming large, up to 35 cm wide, regular rosettes. Lobes 7-20(-30) mm wide, rounded at apex, sinuate at margins, contiguous to slightly overlapping, ivory white to grey-white when dry (turning yellow-brown in the herbarium), green-grey when wet, matt, smooth to warted (pycnidia). Cephalodia globose and partly internal on the lower side, or shrub-like and densely branched on the upper side, the external blue-green-brown or brownish black, up to 2 cm wide. Lower surface white at margins, pale brown over most of the surface, sometimes black in central parts, tomentose, the hairs whitish or brownish 0.15-0.2 mm long, usually with a few, up to 2(-4) mm long, fasciculate, at the base squarrosely branched rhizines. Upper and lower cortex paraplectenchymatous; medulla white. Apothecia rather rare, lecanorine, laminal, sessile and constricted at base, up to 1 cm across, with a concave to flat, glossy brown disc and a smooth, persistent thalline margin. Epithecium brownish; hymenium and hypothecium colourless; paraphyses mostly simple. Asci 8-spored, cylindrical, fissitunicate, the apex of the endoascus with a K/I+ blue ring-shaped structure, Peltigera-type. Ascospores 1-3-septate, hyaline, needle-like, at first hyaline then turning pale yellowish brown, straight to sigmoid, 40-65 x 5-7 μm. Pycnidia laminal, fully immersed into 0.7-1.2 mm wide, conical thalline warts, 0.1-0.2 mm across, the wall 2-layered in upper part, the inner part greyish K+ aeruginose green, the outer part brownish, K-, the ostiole opening as an elongated crack. Conidia bacilliform to rectangular, sometimes swollen at both ends, 3-7 x 1-1.5 μm. Photobiont chlorococcoid in the main thallus, cyanobacterial (Nostoc) in the cephalodia. Spot tests: upper cortex K- or K+ yellow, C-, KC-, P-; medulla K+ pale yellow, C-, KC- or KC+ deep pink, P-. Chemistry: cortex with variable amounts of atranorin (usually in traces); medulla with m-scrobiculin (major) and p-scrobiculin, sometimes substituted by pseudocyphellarin.
Growth form: Foliose, broad lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: very rare
Dry submediterranean belt: extremely rare
Humid submediterranean belt: very rare
Padanian area: absent

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Predictive map

Felix Schumm - CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (19747)
2001/11/26
cephalodia


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (9420)
2001/11/26
apothecia


Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Sardinia, Italy
2014


Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Ricasolia_amplissima.htm
France, 24/03/2014 - Miluccia - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Ricasolia_amplissima.htm
France, 24/03/2014 - Miluccia - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Ricasolia_amplissima.htm
France, 24/03/2014 - Miluccia - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Ricasolia_amplissima.htm
France, 24/03/2014 - Miluccia - Corse


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Col d'Ibardin


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Col d'Ibardin


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Cranou


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Iraty


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Col d'Ibardin


Joel Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Huelgoat


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Cranou


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=363&lang=en
France, Cranou


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Huelgoat
with cephalodia


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Huelgoat
with cephalodia


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Forêt du Cranou
with cephalodia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Col d'Ibardin
with cephalodia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Col d'Ibardin
with cephalodia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Col d'Ibardin
with cephalodia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=983&lang=en
France, Col d'Ibardin
with cephalodia


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Marta Gonzalez Garcia - Centro de Estudios Micologicos Asturianos
Spain, Hayedo del Pico la Cotorra, Soto de Sajambre (León), 10-X-21, 1.400 m, leg & det. Marta González, en madera corticada de Fagus sylvatica, ERD-9244.


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
3052 - Schottland


Felix Schumm - CC BY-SA 4.0
Ki 5017 Portugal - Madeira


Felix Schumm - CC BY-SA 4.0
Ki 5017 Portugal - Madeira


Felix Schumm - CC BY-SA 4.0
Ki 5017 Portugal - Madeira


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bergündeletal Allgäu


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bergündeletal Allgäu


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bergündeletal Allgäu


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bergündeletal Allgäu


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bergündeletal Allgäu


Wolfgang von Brackel - CC BY-4.0
Italy, Calabria


Wolfgang von Brackel - CC BY-4.0
Italy, Calabria, Carlopoli


Wolfgang von Brackel - CC BY-4.0
Italy, Calabria, Carlopoli


Wolfgang von Brackel - CC BY-4.0
Italy, Calabria, Fagnano


Wolfgang von Brackel - CC BY-4.0
Italy, Calabria, Fagnano


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italia, Veneto, in nem. Cansiglio 1855
as Sticta glomerulifera


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italia, Veneto, in nem. Cansiglio 1855
as Sticta glomerulifera


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italia, Veneto, in Alpibus Cadubriae
as Sticta glomerulifera


Source: Schumm F. (2003) Herzogia 16: 91-112 Die Flechtengattung Lobaria auf Madeira.