Athallia holocarpa (Hoffm.) Arup, Frödén & Søchting

in Arup & al., Nord. J. Bot., 31: 36, 2013. Basionym: Verrucaria oblitterata var. holocarpa Hoffm. - Deutsch. Fl., 2, 179, 1796,
Synonyms: Athallia vitellinula ; Callopisma aurantiacum var. holocarpum (Ach.) A. Massal.; Caloplaca aurantiaca var. holocarpa (Ach.) Th. Fr.; Caloplaca holocarpa (Ach.) A.E. Wade; Caloplaca pyracea var. holocarpa (Ach.) Th. Fr.; Caloplaca vitellinula (Nyl.) H. Olivier; Lecanora vitellinula Nyl.; Placodium aurantiacum var. holocarpum (Ach.) Anzi; Placodium pyraceum var. holocarpum (Ach.) Anzi
Description: Thallus absent or developed only around apothecia, sometimes continuous, slightly cracked and verruculose; grey to grey-orange. Cortex usually poorly developed, composed of an amorphous layer or indistinctly paraplectenchymatous. Apothecia frequent, zeorine, 0.3-1(-l.2) mm across, with a flat to slightly convex, yellow-orange to orange disc and a concolorous or slightly paler proper margin, rarely with a poorly developed thalline margin in the basal part of young apothecia. Proper exciple 25-60(-75) µm wide, of irregularly radiating, thick-walled hyphae with rather long and narrow cells, measuring 5-15 x 1-3.5 µm; epithecium orange, granular, K+ purple-red, C-; hymenium colourless, (55-)65-80(-100) μm high, sometimes inspersed with oil droplets; paraphyses simple to sparingly branched in upper part, 2-2.5 μm thick at mid-level, the apical cells to 3-5 μm wide; hypothecium colourless, 50-80(-100) μm high, sometimes inspersed with oil droplets. Asci 8-spored, cylindrical, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, ellipsoid to broadly ellipsoid, (9.5-)10-14(-15.5) x 5-8.5 μm, the equatorial thickening (“septum”) (3-)3.5-5(-5.8) μm, 1/3-1/2 of spore length. Photobiont chlorococcoid. Spot tests: thallus K- or K+ purple-red, C-, KC-, P-; apothecia K+ purple-red, C-. Chemistry: apothecia and orange-pigmented parts of thallus (when present) with parietin (major), fallacinal, emodin, teloschistin and parietinic acid (minor).
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Pioneer species

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: rather rare
Montane belt: rather rare
Dry submediterranean belt: rather rare
Humid submediterranean belt: rare
Padanian area: rare

pH of the substrata:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
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Occurrence data
Predictive map

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29938)
2001/12/03


Curtis Randall Björk- CC BY-SA 4.0
British Columbia, Vancouver Island, Albert Head


Manuel Gil
Spain, Canary Islands, Lanzarote


P.L. Nimis CC BY-SA 4.0
TSB 33408


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A4/Athallia_holocarpa.htm
France, Près du cimetière, Saint-Prejet-Armandon - Haute-Loire
18/11/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A4/Athallia_holocarpa.htm
France, Près du cimetière, Saint-Prejet-Armandon - Haute-Loire
18/11/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A4/Athallia_holocarpa.htm
France, Près du cimetière, Saint-Prejet-Armandon - Haute-Loire
18/11/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A4/Athallia_holocarpa.htm
France, Près du cimetière, Saint-Prejet-Armandon - Haute-Loire
18/11/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A4/Athallia_holocarpa.htm
France, Près du cimetière, Saint-Prejet-Armandon - Haute-Loire
18/11/2015


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A4/Athallia_holocarpa.htm
France, Près du cimetière, Saint-Prejet-Armandon - Haute-Loire
18/11/2015


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=725&lang=en
Belgium, Brest, siliceous rock


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=725&lang=en
Belgium, Brest, siliceous rock


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=725&lang=en
Belgium, Brest, siliceous rock


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=725&lang=en
Belgium, Brest, siliceous rock


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33408)
2001/11/19