Usnea articulata (L.) Hoffm.

Deutschl. Fl., 2: 133, 1796. Basionym: Lichen articulatus L. - Sp. Pl., 2: 1156, 1753.
Synonyms: Usnea articulata subsp. intestiniformis (Ach.) Motyka; Usnea articulata subsp. mediterranea Motyka; Usnea barbata var. articulata (L.) Ach.
Description: Thallus fruticose-filamentous, pale green, long-pendent, to >1 m long, sparingly branched at base, more richly branched towards apices, with a few flexuose lateral branches. Main branches up to 4(-5) mm thick, cylindrical, smooth, the older parts constricted and inflated between constrictions (sausage-like), the surface usually smooth to sparsely papillose, sometimes with scattered comma-like pseudocyphellae. Cortex relatively thin, (5-7 % of the branch diameter); medulla white, very lax and cottony-arachnoid, especially in the inflated parts; central axis white, sometimes visible in the constrictions of old branches. Apothecia unknown in Italian material. Photobiont chlorococcoid. Spot tests: cortex K-, C-, KC-, P-; medulla K- or K+ yellowish slowly turning brown, C-, KC-, P+ red. Chemistry: cortex with usnic acid; medulla with fumarprotocetraric (main) and protocetraric acid (accessory).
Growth form: Fruticose filamentous
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: extremely rare
Dry submediterranean belt: absent
Humid submediterranean belt: extremely rare
Padanian area: absent

pH of the substrata:

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Poleotolerance:

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Predictive map

Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: University of Tartu
Great Britain, Devon, Darmoor National Park, Lydford
2007/04/24
Inflated,sausage-like segments of branches


Andres Saag; Owner: University of Tartu
Great Britain, Devon, Darmoor National Park, Lydford
2006/10/06
Inflated branches


Andres Saag; Owner: University of Tartu
Great Britain, Devon, Darmoor National Park, Lydford
2006/10/06
Pendulous thallus


Andres Saag; Owner: University of Tartu


Andres Saag; Owner: University of Tartu


Andres Saag; Owner: University of Tartu


Andres Saag; Owner: University of Tartu


Andres Saag; Owner: University of Tartu


Harry Taylor; Owner: Natural History Museum London


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Pier Luigi Nimis
Slovenia, Sicily, Trapani, Erice, near the castle
12.2016


Felix Schumm - CC BY-SA 4.0
[2710], Apanien, Kanarische Inseln, Gran Canaria, 7 km südlich von Lanzarote, Felswand an der Straße nach Tejeda oberhalb von Cuevas del Corcho, Nebelzone, ca. 1300 m Schumm 03.01.1978


Felix Schumm - CC BY-SA 4.0
[2710], Apanien, Kanarische Inseln, Gran Canaria, 7 km südlich von Lanzarote, Felswand an der Straße nach Tejeda oberhalb von Cuevas del Corcho, Nebelzone, ca. 1300 m Schumm 03.01.1978


Felix Schumm - CC BY-SA 4.0
[2710], Apanien, Kanarische Inseln, Gran Canaria, 7 km südlich von Lanzarote, Felswand an der Straße nach Tejeda oberhalb von Cuevas del Corcho, Nebelzone, ca. 1300 m Schumm 03.01.1978


Felix Schumm - CC BY-SA 4.0
[2710], Apanien, Kanarische Inseln, Gran Canaria, 7 km südlich von Lanzarote, Felswand an der Straße nach Tejeda oberhalb von Cuevas del Corcho, Nebelzone, ca. 1300 m Schumm 03.01.1978


Felix Schumm - CC BY-SA 4.0
[2710], Apanien, Kanarische Inseln, Gran Canaria, 7 km südlich von Lanzarote, Felswand an der Straße nach Tejeda oberhalb von Cuevas del Corcho, Nebelzone, ca. 1300 m Schumm 03.01.1978


Felix Schumm - CC BY-SA 4.0
[2710], Apanien, Kanarische Inseln, Gran Canaria, 7 km südlich von Lanzarote, Felswand an der Straße nach Tejeda oberhalb von Cuevas del Corcho, Nebelzone, ca. 1300 m Schumm 03.01.1978


Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Usnea_articulata.htm
France, 9/10/2010 - sous le col de Vergio D84, alt : 1300 m - Corse -


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Usnea_articulata.htm
France, 9/10/2010 - sous le col de Vergio D84, alt : 1300 m - Corse -


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Brasparts


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Brennilis


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Cranou


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Cranou


Joel Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, St. Herbot


Joel Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, St. Herbot


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, St. Herbot


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Brennilis


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Cranou


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Cranou


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Commana


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
France, Sant Mikel


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=323&lang=en
Spain, Canary Islands, Tenerife, Teide