Verrucaria maculiformis Kremp.

Flora, 41: 303, 1858, nom. illegit. non Hoffm.
Synonyms: Involucrothele maculiformis (Servít) Servít; Thelidium maculiforme Servít
Description: Thallus crustose, episubstratic, 100-150 μm thick, granular-areolate or fleck-like, subgelatinous when wet, medium to dark brown, glossy, smooth to rarely minutely uneven, sometimes delimited by an inconspicuous, brownish prothallus. Cortex thin, the cells with brownish-pigmented walls; algal layer continuous, 20-40 μm thick, the algal cells surrounded by paraplectenchymatous hyphae; medulla loose, paraplectenchymatous, with crystals visible under polarized light, without a black basal layer. Perithecia half to 1/4 immersed in the thallus, forming moderate, 0.15-0.25 mm wide projections, sometimes with a very thin thalline layer in lower part. Involucrellum limited to the upper half of exciple, conical, dark brown, 30-50 μm thick; exciple 0.15-0.2 mm across, the wall at first colourless, then brown; hamathecium of thick, unbranched, 15-20 μm long periphyses and periphysoids, interascal filaments absent; hymenial gel hemiamyloid, I+ red (I+ blue at very low concentrations of I), K/I+ blue. Asci 8-spored, clavate, I-, fissitunicate, the wall thickened above, with an ocular chamber, dehiscent by extrusion of an endotunica to form a delicate rostrum, Verrucaria-type. Ascospores 1-celled, hyaline, ellipsoid, 14-20 x 6-9 μm. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Poorly known taxon in need of further study

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: extremely rare
Dry submediterranean belt: very rare
Humid submediterranean belt: very rare
Padanian area: absent

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Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_V/Verrucaria_maculiformis.htm
France, sur galet siliceux - Miribel-Jonage - 28/06/2011


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_V/Verrucaria_maculiformis.htm
France, sur galet siliceux - Miribel-Jonage - 28/06/2011


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_V/Verrucaria_maculiformis.htm
France, sur galet siliceux - Miribel-Jonage - 28/06/2011


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_V/Verrucaria_maculiformis.htm
France, sur galet siliceux - Miribel-Jonage - 28/06/2011


Zschacke, H. (1934) Epigloeaceae, Verrucariaceae und Dermatocarpaceae. In: Dr. L. Rabenhorst‘s Kryptogamen-Flora, Band 9, Abt. 1, Teil 1. Akademische Verlagsgesellschaft, Leipzig, 695 pp. - Public Domain