Brodoa intestiniformis (Vill.) Goward

Bryologist, 89: 222, 1986. Basionym: Lichen intestiniformis Vill. - Hist. Pl. Dauphiné, 3: 947, 1789.
Synonyms: Hypogymnia encausta (Sm.) Walt. Watson; Hypogymnia intestiniformis (Vill.) Räsänen; Imbricaria encausta (Sm.) DC.; Menegazzia encausta (Sm.) Navàs; Parmelia ceratophylla var. multipuncta (Ehrh.) Schaer.; Parmelia encausta (Sm.) Nyl.; Parmelia encausta var. multipuncta (Ehrh.) Th. Fr.; Parmelia intestiniformis (Vill.) Ach.
Description: Thallus foliose, heteromerous, dorvisentral, adnate, forming 2-5 cm wide, orbicular or irregular rosettes. Lobes up to c. 2.5 cm long, 0.5-1(-2) mm broad, more or less convex, stiff, contiguous or overlapping; center of thallus with narrower (0.2-0.4 mm), flattened secondary lobules which arise perpendicularly to the surface. Upper surface pale grey to brownish, darkening or blackening especially on the lobules and towards the tips, and there becoming glossy, reticulate-maculate at least in the apical 2-3 mm. Lower surface black, brown toward the tips, smooth to weakly and shallowly wrinkled, erhizinate, with scattered hapters. Upper cortex paraplectenchymatous, the cell walls with Cetraria-type lichenan, with a non-pored epicortex; medulla white, solid; lower cortex paraplectenchymatous, without an epicortex. Apothecia rare, lecanorine, up to 5(-7) mm across, with a dark brown disc and a thin, entire to sublobulate thalline margin. Epithecium brownish; hypothecium and hymenium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, subglobose to broadly ellipsoid, 10-12 x 6-8 µm. Pycnidia numerous, immersed, the visible part black to grey-brown. Conidia bifusiform, 5-7 x c. 1 µm. Photobiont chlorococcoid. Spot tests: upper cortex K+ yellow, C-, KC-, P-; medulla K+ slowly reddish to orange-brown, C-, KC-, P+ orange, UV-. Chemistry: upper cortex with atranorin and chloroatranorin; medulla with fumarprotocetraric acid.
Growth form: Foliose, narrow lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather common
Subalpine belt: common
Montane belt: absent
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent

pH of the substrata:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (12970)
2001/11/22
central lobuli


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35222)
2002/07/12
central lobules


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (12970)
2001/11/22
detail of marginal lobes


Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)


Juri Nascimbene - CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Val di Rabbi
2006


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Pier Luigi Nimis CC BY-SA 4.0
TSB 41693


Pier Luigi Nimis - CC BY-SA 4.0
TSB 37046


Pier Luigi Nimis - CC BY-SA 4.0
TSB 37046


Pier Luigi Nimis - CC BY-SA 4.0
TSB 37046


Pier Luigi Nimis - CC BY-SA 4.0
TSB 37047


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_B/Brodoa_intestiniformis.htm
France, Mont Cenis - 2081 m - Savoie
4/8/2011


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_B/Brodoa_intestiniformis.htm
France, Mont Cenis - 2081 m - Savoie
4/8/2011


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_B/Brodoa_intestiniformis.htm
France, Mont Cenis - 2081 m - Savoie
4/8/2011


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
On acid rocks. Austria: Alps.


Wolfgang von Brackel - CC BY-4.0
Czech Republic, Stribro


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bayerwald, Lusen


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bayerwald Plöckenstein


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bayerwald Plöckenstein


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Bayerwald Plöckenstein


Wolfgang von Brackel - CC BY-4.0
Greenland, Blomsterdalen


Wolfgang von Brackel - CC BY-4.0
Greenland, Tugtutoq


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, in Al. Cadubriae
as Parmelia encausta


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, in Alp. Cadubriae M. Spina 1855
as Parmelia encausta