Calogaya pusilla (A. Massal.) Arup, Frödén & Søchting | Dolichens

Calogaya pusilla (A. Massal.) Arup, Frödén & Søchting

in Arup & al., Nord. J. Bot., 31: 39, 2013. Basionym: Physcia pusilla A. Massal. - Atti Ist. Ven., ser. 2, vol. III, append. III: 59, fig. VII, 1852.

Synonyms: Caloplaca murorum f. pulvinata (A. Massal.) Ozenda & Clauzade; Caloplaca murorum var. pulvinata (A. Massal.) Jatta; Caloplaca pusilla (A. Massal.) Zahlbr.; Caloplaca saxicola subsp. pulvinata (A. Massal.) Clauzade & Cl. Roux; Gasparrinia pusilla (A. Massal.) Tornab.; Physcia murorum var. pulvinata A. Massal.; Physcia murorum var. pulvinata f. euphora A. Massal.; Physcia pusilla var. turgida A. Massal.; Placodium pusillum (A. Massal.) Anzi

Description: Thallus crustose-placodioid, episubstratic, salmon-coloured to brownish orange, often thinly white pruinose, especially in central part, forming regular, up to 1.8 cm wide rosettes, several thalli often merging to cover larger surfaces. Central part areolate, with flat to convex, 0.3-1.3 mm wide areoles formed from lobe fragmentation; marginal lobes convex, sometimes becoming flat toward the somewhat broader apex, finger-like or strongly branched, always contiguous, 0.3-1.7 mm long. 0.2-1.2 mm wide. Thallus 100-365 μm thick; upper cortex 14-65(-85) μm thick, with an orange, K+ purple-red upper layer and a colourless inner layer, scleroplectenchymatous, the upper part with many crystals visible under polarized light; medulla quite lax. Apothecia pseudolecanorine or zeorine, abundant, crowded, 0.2-1.2 mm across, rounded or deformed by mutual compression, usually on areoles and lobe bases. Disk flat to strongly convex, orange to brownish orange, slightly pruinose, smooth; margin in pseudolecanorine apothecia 30-130 μm thick, concolorous with disc; in zeorine apothecia, proper margin 30-50 μm thick and thalline margin 30-130 μm thick, the former slightly lighter than disk, not prominent, the latter pruinose. Epithecium (6-)8-10(-12) μm thick, orange, K+ purple-red; hymenium colourless, 50-80(-95) μm high; paraphyses septate, forked or weakly branched subapically, (1.5-)2-3(-3.5) μm at base, the apical cells (2.5-)3-7(-8.5) μm wide; hypothecium colourless, sclero-prosoplectenchymatous. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, broadly ellipsoid (9-)10.5-13.5(-16) x (4.5-)5.5-7(-8.7) μm, the equatorial thickening (“septum”) (2.5-)3-4.5(-6) μm, > 1/3 of spore length. Photobiont chlorococcoid. Spot tests: thallus and apothecia K+ purple-red, C-, KC-, P-. Chemistry: thallus and apothecia with parietin (major), fallacinal, emodin, teloschistin and parietinic acid (minor), corresponding with chemosyndrome A of Søchting (1997).

Growth form: Crustose placodiomorph

Substrata: rocks

Photobiont: green algae other than Trentepohlia

Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rare

Subalpine belt: rather rare

Montane belt: rather common

Dry submediterranean belt: very common

Humid submediterranean belt: very common

Padanian area: rather rare

pH of the substrata:

1

2

3

4

5

Solar irradiation:

1

2

3

4

5

Aridity:

1

2

3

4

5

Eutrophication:

1

2

3

4

5

Poleotolerance:

0

1

2

3

Altitudinal distribution:

1

2

3

4

5

6

Rarity

absent

extremely rare

very rare

rare

rather rare

rather common

common

very common

extremely common

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Occurrence data

Predictive map

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder

On a calcareous wall. Central Europe; Germany: Hesse.

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (11201)
2001/12/07

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (3707)
2001/11/19

Antonio Valero; Owner: Antonio Valero

Spain
2010

Antonio Valero; Owner: Antonio Valero

Spain
2010

Antonio Valero; Owner: Antonio Valero

Spain
2010

Antonio Valero; Owner: Antonio Valero

Spain
2010

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Triedte

Italy, Veneto, Venezia, Venice, Giardini Biennale
07/2016

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Triedte

Italy, Veneto, Venezia, Venice, Giardini Biennale
07/2016

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Triedte

Italy, Veneto, Venezia, Venice, Giardini Biennale
07/2016

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Triedte

Italy, Veneto, Venezia, Venice, Giardini Biennale
07/2016

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Triedte

Italy, Veneto, Venezia, Venice, Giardini Biennale
07/2016

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Triedte

Italy, Veneto, Venezia, Venice, Giardini Biennale
07/2016

Andrea Moro; Owner: Department of Life Sciences, University of Triedte

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017

Felix Schumm – CC BY-SA 4.0

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Felix Schumm CC BY-SA 4.0

[16565], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Göppingen. Messelstein bei Donzdorf, an Kalkfelsen, 48.69045° N, 9.83113° E, 750 m. Leg. et det. Schumm 25.07.2010.
as Caloplaca saxicola

Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.

as Caloplaca murorum

Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0

TSB 37183

Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0

TSB 37183

Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/

Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/

Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/

Austria, Schoeckl-Sendermast, auf Gedenkstein

Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/

Austria, Schoeckl-Sendermast, auf Gedenkstein

Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Calogaya_pusilla.htm

France, session AFL 2014 - Col de la Bonnette -

Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Calogaya_pusilla.htm

France, session AFL 2014 - Col de la Bonnette -

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (33347)
2001/12/03
detail of marginal lobes

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder

SE-Europe; N-Cyprus; E of Girne; Beşparmak Mountains; SE of Kaplica; Kantara Castle

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder

SE-Europe; N-Cyprus; E of Girne; Beşparmak Mountains; SE of Kaplica; Kantara Castle

Source: https://nhm2.uio.no/typephotos/lichens/typephotoS.php?f=O-L-110841d_s.jpg

ITALY: [s. loc.], ., ISOTYPE

Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC

Italy, Veneto, in op. Velo 1855
as Physcia murorum pulvinata

Bologna

Biome

Pon

Trieste

Trieste