Cladonia arbuscula (Wallr.) Flot.
in Wendt, Therm. Warmbrunn: 94, 1839. Basionym: Patellaria foliacea var. arbuscula Wallr. - Naturgesch. Säulchen-Flecht.: 169, 1829.
Synonyms: Cladina arbuscula (Wallr.) Hale & W.L. Culb.; Cladina arbuscula subsp. squarrosa (Wallr.) Burgaz; Cladonia arbuscula subsp. squarrosa (Wallr.) Ruoss; Cladonia sylvatica auct.; Patellaria arbuscula (Wallr.) Wallr.; Patellaria coccinea var. squarrosa Wallr.
Description: Primary thallus crustose, ephemeral and rarely seen, consisting of subglobular, 0.1-1.5 mm wide warts. Podetia densely anisotomically (i.e. main branches thicker) mainly tri- or tetrachotomously branched, shrub-like, hollow inside, esquamulose, ecorticate, the surface compact or finely arachnoid, 4-10 cm tall, loosely attached, dying from the base, pale yellowish white, with open axils, the necrotic parts yellowish white; main axis distinct, 0.7-1.5(-2) mm thick; terminal branchlets unilaterally deflexed, curved, often browned at tips. Apothecia rare, terminal, dark brown, convex. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed on the tips of podetia; pycnidial jelly colourless. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K- or K+ yellowish slowly turning brown, C-, KC+ yellow, P+ orange-red (rarely P+ deep yellow or rarely P-), UV-. Chemistry: the most common chemotype contains usnic and isousnic acids (major) and substances of the fumarprotocetraric complex; other, much rarer chemotypes have no fumarprotocetraric acid (P-), or additional psoromic acid (P+ yellow).
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation
Commonnes-rarity: (info)
Alpine belt: rather rare
Subalpine belt: common
Montane belt: extremely rare
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Current prediction (1981-2010)
Future prediction (2071-2100) SSP 1-2.6
Future prediction (2071-2100) SSP 5-8.5Predictive maps according to Francesconi et al. 2025
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (25106)
2003/02/15

Felix Schumm- CC BY-SA 4.0
[19513], Schweiz, Wallis, Champex westl. Orsières, Val de Arpette, Granituntergrund, ca. 1700 m. Leg. et det. F. Schumm 22.08.1996 (SVBL-Exkursion).

Felix Schumm- CC BY-SA 4.0
[19513], Schweiz, Wallis, Champex westl. Orsières, Val de Arpette, Granituntergrund, ca. 1700 m. Leg. et det. F. Schumm 22.08.1996 (SVBL-Exkursion).

Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
September 2017

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.
with C. rangiferina (left)

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation
Commonnes-rarity: (info)
Alpine belt: rather rare
Subalpine belt: common
Montane belt: extremely rare
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent
pH of the substrata:
1 | 2 | 3 | 4 | 5 |
Solar irradiation:
1 | 2 | 3 | 4 | 5 |
Aridity:
1 | 2 | 3 | 4 | 5 |
Eutrophication:
1 | 2 | 3 | 4 | 5 |
Poleotolerance:
0 | 1 | 2 | 3 |
Altitudinal distribution:
1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map



Predictive maps according to Francesconi et al. 2025

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (25106)
2003/02/15

Felix Schumm- CC BY-SA 4.0
[19513], Schweiz, Wallis, Champex westl. Orsières, Val de Arpette, Granituntergrund, ca. 1700 m. Leg. et det. F. Schumm 22.08.1996 (SVBL-Exkursion).

Felix Schumm- CC BY-SA 4.0
[19513], Schweiz, Wallis, Champex westl. Orsières, Val de Arpette, Granituntergrund, ca. 1700 m. Leg. et det. F. Schumm 22.08.1996 (SVBL-Exkursion).

Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
September 2017

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.
with C. rangiferina (left)

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.
