Cladonia arbuscula (Wallr.) Flot.

in Wendt, Therm. Warmbrunn: 94, 1839. Basionym: Patellaria foliacea var. arbuscula Wallr. - Naturgesch. Säulchen-Flecht.: 169, 1829.
Synonyms: Cladina arbuscula (Wallr.) Hale & W.L. Culb.; Cladina arbuscula subsp. squarrosa (Wallr.) Burgaz; Cladonia arbuscula subsp. squarrosa (Wallr.) Ruoss; Cladonia sylvatica auct.; Patellaria arbuscula (Wallr.) Wallr.; Patellaria coccinea var. squarrosa Wallr.
Description: Primary thallus crustose, ephemeral and rarely seen, consisting of subglobular, 0.1-1.5 mm wide warts. Podetia densely anisotomically (i.e. main branches thicker) mainly tri- or tetrachotomously branched, shrub-like, hollow inside, esquamulose, ecorticate, the surface compact or finely arachnoid, 4-10 cm tall, loosely attached, dying from the base, pale yellowish white, with open axils, the necrotic parts yellowish white; main axis distinct, 0.7-1.5(-2) mm thick; terminal branchlets unilaterally deflexed, curved, often browned at tips. Apothecia rare, terminal, dark brown, convex. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed on the tips of podetia; pycnidial jelly colourless. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K- or K+ yellowish slowly turning brown, C-, KC+ yellow, P+ orange-red (rarely P+ deep yellow or rarely P-), UV-. Chemistry: the most common chemotype contains usnic and isousnic acids (major) and substances of the fumarprotocetraric complex; other, much rarer chemotypes have no fumarprotocetraric acid (P-), or additional psoromic acid (P+ yellow).
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: common
Montane belt: extremely rare
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php - Courtesy: Anita Stridvall


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (25106)
2003/02/15


Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)


Giancarlo Medici; Owner: Giancarlo Medici
Sweden, Svezia
2010


Felix Schumm- CC BY-SA 4.0
[19513], Schweiz, Wallis, Champex westl. Orsières, Val de Arpette, Granituntergrund, ca. 1700 m. Leg. et det. F. Schumm 22.08.1996 (SVBL-Exkursion).


Felix Schumm- CC BY-SA 4.0
[19513], Schweiz, Wallis, Champex westl. Orsières, Val de Arpette, Granituntergrund, ca. 1700 m. Leg. et det. F. Schumm 22.08.1996 (SVBL-Exkursion).


Juri Nascimbene - CC BY-SA 4.0
Paneveggio, Reg. Trentino-Alto Adige Prov. Trento, Italy.
2009


Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
September 2017


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.
with C. rangiferina (left)


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Faedo (Quirós-Asturias), 29-VIII-2023, sobre un talud rocoso ácido, en zonas umbrías, leg. M. González & det. Helena Rodríguez e Isaac Rubio, conf. A. R. Burgaz, MGG-46.


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Allgäu, Zeiger-Südhang


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Büchelbach-Bayerwald


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Erbendorf


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Föhrenbühl -


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Neumühlschlag


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Tennenlohe


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Tennenlohe


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Tennenlohe


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Tennenlohe


Triin Aimla; Owner: University of Tartu
originally as Cladina arbuscula