Cladonia foliacea (Huds.) Willd.

Fl. Berol.: 363, 1787. Basionym: Lichen foliaceus Huds. - Fl. Angl.: 457, 1762
Synonyms: Cladonia alcicornis (Lightf.) Fr.; Cladonia convoluta (Lam.) Anders; Cladonia endiviifolia auct. p.p.; Cladonia foliacea f. convoluta (Lam.); Cladonia foliacea subsp. convoluta (Lam.) Cretz.; Cladonia foliacea var. alcicornis (Lightf.) Schaer.; Cladonia foliacea var. convoluta (Lam.) Vain.
Description: Primary thallus squamulose, the squamules large, 6-40 mm long, 2-20 mm wide, sometimes with black or white hairs at the margins, persistent, forming more or less dense, 4-10 cm broad mats, the upper surface yellowish green to greenish-grey above, the lower surface cream-coloured or more rarely white, finely arachnoid. Podetia very rare, hollow inside, up to 1.5(-2) cm tall, with irregular cups, the surface corticate, smooth, very rarely with squamules on the margin of cups. Apothecia pale brown, convex, often confluent. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia frequent on the primary squamules or at the margins of cups, sessile to prominent, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K- or K+ yellowish slowly turning brown, C-, KC+ yellowish, P+ red, UV-. Chemistry: fumarprotocetraric and usnic acids, rarely with additional psoromic acid, frequently with traces of zeorin.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: rather rare
Dry submediterranean belt: very common
Humid submediterranean belt: very common
Padanian area: extremely rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
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extremely common
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

Felix Schumm – CC BY-SA 4.0
[9372], Spanien, La Gomera, bei Arure an der Straße nach Las Hayas. Trockener Abhang mit Felsblöcken und Euphorbia. 28°07.895 N, 17°18.983 W, 870 m. Leg. Schumm 14.02.2002.


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php - Courtesy: Anita Stridvall
Calcicolous form (f. convoluta)


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (4051)
2001/12/04


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10122)
2001/12/04


Andres Saag; Owner: University of Tartu


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10122)
2001/12/04
Calcicolous form (f. convoluta)


A. Moro; Owner: Department of Life Sciences, University of Trieste
Sicilia, TP, Segesta, presso i templi
2008.04.05
Calcicolous form (f. convoluta)


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Gabriele Gheza, Tromello (PV)
Calcicolous form (f. convoluta)


Felix Schumm – CC BY-SA 4.0
[9372], Spanien, La Gomera, bei Arure an der Straße nach Las Hayas. Trockener Abhang mit Felsblöcken und Euphorbia. 28°07.895 N, 17°18.983 W, 870 m. Leg. Schumm 14.02.2002.,
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start, 4: Position von Norstictinsäure, 7: Position von Atranorin, us: Usninsäure fpr: Fumarprotocetrarsäure pr: Protocetrarsäure


Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Parco Nazionale delle Foreste Casentinesi
2010
Calcicolous form (f. convoluta)


Simonetta Peruzzi
Italy, Toscana, Parco Nazionale delle Foreste Casentinesi
2010
Calcicolous form (f. convoluta)


Domenico Puntillo - CC BY-SA 4.0
Italy, Calabria
Calcicolous form (f. convoluta)


Felix Schumm – CC BY-SA 4.0
[12537], Griechenland, W-Samos, Mt. Kerketeas, beim Kloster Perados, 37°44.926'N, 26°38.888'E, 575 m. Leg. Schumm, Düll & Wunder 13.05.2006, det. Schumm.
Calcicolous form (f. convoluta)


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Milano, Magenta, valle del Ticino, 116 m
17/05/2019


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Pavia, Vigevano, valle del Ticino, 75 m
09/2018


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Pavia, Dossi della Lomellina, 115 m
03/2011


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta


Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta


Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta


Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Hainberg


Wolfgang von Brackel - CC BY-4.0
Romania, Mihail Kogalniceanu


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, L. Abano
as C. endiviaefolia