Cladonia foliacea (Huds.) Willd.
Fl. Berol.: 363, 1787. Basionym: Lichen foliaceus Huds. - Fl. Angl.: 457, 1762
Synonyms: Cladonia alcicornis (Lightf.) Fr.; Cladonia convoluta (Lam.) Anders; Cladonia endiviifolia auct. p.p.; Cladonia foliacea f. convoluta (Lam.); Cladonia foliacea subsp. convoluta (Lam.) Cretz.; Cladonia foliacea var. alcicornis (Lightf.) Schaer.; Cladonia foliacea var. convoluta (Lam.) Vain.
Description: Primary thallus squamulose, the squamules large, 6-40 mm long, 2-20 mm wide, sometimes with black or white hairs at the margins, persistent, forming more or less dense, 4-10 cm broad mats, the upper surface yellowish green to greenish-grey above, the lower surface cream-coloured or more rarely white, finely arachnoid. Podetia very rare, hollow inside, up to 1.5(-2) cm tall, with irregular cups, the surface corticate, smooth, very rarely with squamules on the margin of cups. Apothecia pale brown, convex, often confluent. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia frequent on the primary squamules or at the margins of cups, sessile to prominent, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K- or K+ yellowish slowly turning brown, C-, KC+ yellowish, P+ red, UV-. Chemistry: fumarprotocetraric and usnic acids, rarely with additional psoromic acid, frequently with traces of zeorin.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Montane belt: rather rare
Dry submediterranean belt: very common
Humid submediterranean belt: very common
Padanian area: extremely rare
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Current prediction (1981-2010)
Future prediction (2071-2100) SSP 1-2.6
Future prediction (2071-2100) SSP 5-8.5Predictive maps according to Francesconi et al. 2025
Felix Schumm – CC BY-SA 4.0
[9372], Spanien, La Gomera, bei Arure an der Straße nach Las Hayas. Trockener Abhang mit Felsblöcken und Euphorbia. 28°07.895 N, 17°18.983 W, 870 m. Leg. Schumm 14.02.2002.

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php - Courtesy: Anita Stridvall
Calcicolous form (f. convoluta)

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (4051)
2001/12/04

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10122)
2001/12/04

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10122)
2001/12/04
Calcicolous form (f. convoluta)

A. Moro; Owner: Department of Life Sciences, University of Trieste
Sicilia, TP, Segesta, presso i templi
2008.04.05
Calcicolous form (f. convoluta)

Felix Schumm – CC BY-SA 4.0
[9372], Spanien, La Gomera, bei Arure an der Straße nach Las Hayas. Trockener Abhang mit Felsblöcken und Euphorbia. 28°07.895 N, 17°18.983 W, 870 m. Leg. Schumm 14.02.2002.,
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start, 4: Position von Norstictinsäure, 7: Position von Atranorin, us: Usninsäure fpr: Fumarprotocetrarsäure pr: Protocetrarsäure

Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Parco Nazionale delle Foreste Casentinesi
2010
Calcicolous form (f. convoluta)

Simonetta Peruzzi
Italy, Toscana, Parco Nazionale delle Foreste Casentinesi
2010
Calcicolous form (f. convoluta)

Felix Schumm – CC BY-SA 4.0
[12537], Griechenland, W-Samos, Mt. Kerketeas, beim Kloster Perados, 37°44.926'N, 26°38.888'E, 575 m. Leg. Schumm, Düll & Wunder 13.05.2006, det. Schumm.
Calcicolous form (f. convoluta)

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Milano, Magenta, valle del Ticino, 116 m
17/05/2019

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Pavia, Vigevano, valle del Ticino, 75 m
09/2018

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Pavia, Dossi della Lomellina, 115 m
03/2011

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta

Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta

Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Montane belt: rather rare
Dry submediterranean belt: very common
Humid submediterranean belt: very common
Padanian area: extremely rare
pH of the substrata:
1 | 2 | 3 | 4 | 5 |
Solar irradiation:
1 | 2 | 3 | 4 | 5 |
Aridity:
1 | 2 | 3 | 4 | 5 |
Eutrophication:
1 | 2 | 3 | 4 | 5 |
Poleotolerance:
0 | 1 | 2 | 3 |
Altitudinal distribution:
1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map



Predictive maps according to Francesconi et al. 2025

Felix Schumm – CC BY-SA 4.0
[9372], Spanien, La Gomera, bei Arure an der Straße nach Las Hayas. Trockener Abhang mit Felsblöcken und Euphorbia. 28°07.895 N, 17°18.983 W, 870 m. Leg. Schumm 14.02.2002.

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php - Courtesy: Anita Stridvall
Calcicolous form (f. convoluta)

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (4051)
2001/12/04

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10122)
2001/12/04

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10122)
2001/12/04
Calcicolous form (f. convoluta)

A. Moro; Owner: Department of Life Sciences, University of Trieste
Sicilia, TP, Segesta, presso i templi
2008.04.05
Calcicolous form (f. convoluta)

Felix Schumm – CC BY-SA 4.0
[9372], Spanien, La Gomera, bei Arure an der Straße nach Las Hayas. Trockener Abhang mit Felsblöcken und Euphorbia. 28°07.895 N, 17°18.983 W, 870 m. Leg. Schumm 14.02.2002.,
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start, 4: Position von Norstictinsäure, 7: Position von Atranorin, us: Usninsäure fpr: Fumarprotocetrarsäure pr: Protocetrarsäure

Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Parco Nazionale delle Foreste Casentinesi
2010
Calcicolous form (f. convoluta)

Simonetta Peruzzi
Italy, Toscana, Parco Nazionale delle Foreste Casentinesi
2010
Calcicolous form (f. convoluta)

Felix Schumm – CC BY-SA 4.0
[12537], Griechenland, W-Samos, Mt. Kerketeas, beim Kloster Perados, 37°44.926'N, 26°38.888'E, 575 m. Leg. Schumm, Düll & Wunder 13.05.2006, det. Schumm.
Calcicolous form (f. convoluta)

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Toscana, Firenze, Fiesole, Monte Ceceri, 390 m
31/01/2020

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Milano, Magenta, valle del Ticino, 116 m
17/05/2019

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 600 m
03/2017

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Pavia, Vigevano, valle del Ticino, 75 m
09/2018

Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Pavia, Dossi della Lomellina, 115 m
03/2011

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Provence, Manosque
as Cladonia convoluta

Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta

Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta

Arnaud Delhoume- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=745&lang=en
France, Nievre
as Cladonia convoluta

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=148&lang=en
France, Côte d'Opale

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Petralia Sottana (Palermo-Sicilia), 6-IV-2018, sobre arenisca, a 1.124 m de altitud, leg. & det. A. R. Burgaz, MGG-105.
