Thyrea confusa Henssen

in Henssen & Jørgensen, Lichenologist, 22: 146, 1990.
Synonyms: Omphalaria pulvinata auct. non (Schaer.) Nyl.; Thyrea pulvinata auct. non (Schaer.) A. Massal.
Description: Thallus foliose to subfruticose, polyphyllous, gelatinous when wet, forming small cushions or up to 2 cm wide rosettes. Squamules 2-5 mm wide, to 1 cm long, erect or ascending, flat or weakly channelled on lower surface, irregularly branched, broadly rounded at apex, black but most often bluish-grey pruinose, often with small, globose, granular isidia, otherwise smooth; lower surface plicate, often paler than upper surface. Thallus ecorticate, 250-500 µm thick; medulla 200-300 µm thick, of irregularly arranged, lax hyphae, in the upper and lower parts consisting of mainly anticlinally oriented hyphae enclosing the photobiont cells. Apothecia rare, immersed or semi-immersed, hemiangiocarpic, forming pycnoascocarps from ascogonia beneath pycnidia, small, the disc punctiform, slightly depressed, brownish, with a persisting thalline margin. Proper exciple colourless, up to 10 µm wide; epithecium poorly differentiated; hymenium colourless, up to 150 µm high, K/I+ blue; paraphyses distinctly septate, sparingly branched and anastomosing, the apical cells slightly thickened, up to 3 µm wide; hypothecium colourless. Asci 8-spored, prototunicate, cylindrical-clavate, thin-walled, with an external, I+ blue cap. Ascospores 1-celled, hyaline, globose to broadly ellipsoid, thin-walled, (7.5-)8-12(-15) x 5-7.5(-10) µm. Pycnidia immersed, globose, unilocular, 90-120 µm across. Conidia cylindrical, 2-3 x 1-1.5 µm. Photobiont cyanobacterial, chroococcoid, with a few cells measuring (4-)6-8(-10) µm, surrounded by a brownish gelatinous sheath. Spot test: all negative. Chemistry: without lichen substances.
Growth form: Fruticose

Substrata: rocks
Photobiont: cyanobacteria, coccaceous (e.g. Gloeocapsa)
Reproductive strategy: mainly sexual
On otherwise dry surfaces with short periods of water seepage after rain

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: extremely rare
Montane belt: very rare
Dry submediterranean belt: rather rare
Humid submediterranean belt: rare
Padanian area: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
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rather rare
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common
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Occurrence data
Predictive map

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (2294)
2001/12/11


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_T/Textes_T2/Thyrea_confusa.htm
France, 18/5/2015 - Session AFL en Dordogne - Bourzolles


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_T/Textes_T2/Thyrea_confusa.htm
France, 18/5/2015 - Session AFL en Dordogne - Bourzolles


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_T/Textes_T2/Thyrea_confusa.htm
France, 18/5/2015 - Session AFL en Dordogne - Bourzolles


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_T/Textes_T2/Thyrea_confusa.htm
France, 18/5/2015 - Session AFL en Dordogne - Bourzolles


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_T/Textes_T2/Thyrea_confusa.htm
France, 18/5/2015 - Session AFL en Dordogne - Bourzolles


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL56996], China, Yunnan Prov., Yunnan Co., Stone Forest, Parkland with limestone outcrops, c.. 2000 m, 24°49’01’’ N, 103°19’27’’ E, on limestone. Leg. A. Aptroot (no 56996), 27.10.2002, det. A. Aptroot, 2003, conf. M. Schulz, 2005.


Felix Schumm - CC BY-SA 4.0
[ABL77834], Brazil, Mato Grosso do Sul, Bodoquena, Dente de Cão, summit, in Atlantic rain forest on exposed limestone. 20°47’05’’ S, 56°45’0’’ W, 450 m. Leg. A. Aptroot (no 77834), 7.11.2018, det. A. Aptroot, 2019.


Felix Schumm - CC BY-SA 4.0
[ABL77834], Brazil, Mato Grosso do Sul, Bodoquena, Dente de Cão, summit, in Atlantic rain forest on exposed limestone. 20°47’05’’ S, 56°45’0’’ W, 450 m. Leg. A. Aptroot (no 77834), 7.11.2018, det. A. Aptroot, 2019.


Felix Schumm - CC BY-SA 4.0
[ABL77834], Brazil, Mato Grosso do Sul, Bodoquena, Dente de Cão, summit, in Atlantic rain forest on exposed limestone. 20°47’05’’ S, 56°45’0’’ W, 450 m. Leg. A. Aptroot (no 77834), 7.11.2018, det. A. Aptroot, 2019.


Felix Schumm - CC BY-SA 4.0
[ABL77834], Brazil, Mato Grosso do Sul, Bodoquena, Dente de Cão, summit, in Atlantic rain forest on exposed limestone. 20°47’05’’ S, 56°45’0’’ W, 450 m. Leg. A. Aptroot (no 77834), 7.11.2018, det. A. Aptroot, 2019.


Detail from: https://gzu.jacq.org/GZU000291949
GZU000291949 - Collector Clauzade,G. & Roux,C. Vězda, A.: Lichenes Selecti Exsiccati 1157 Date 1972-12-07 Location France / Languedoc-Roussillon Label Gallia. Hérault. Mireval, Creux de Miege ; Alt. 50 m Habitat Ad saxa calcarea