Cladonia macroceras (Delise) Hav.

Bergens Mus. ?rbok, Naturvid. Rekke, 1927, 3: 12, 1928. Basionym: Cenomyce gracilis var. macroceras Delise in Duby - Bot. Gall., 2: 624, 1830.
Synonyms: Cladonia elongata auct. p.p.; Cladonia gracilis var. macroceras (Delise) Flot.
Description: Primary thallus squamulose, often ephemeral but sometimes persistent and forming a bullate crust, the squamules middle-sized (1-4 mm), thick, esorediate, grey green to brown above, white beneath. Podetia robust and elongate, hollow inside, brownish grey to dark brown, becoming reddish brown to almost black at base, up to 15 cm tall and 1-4(-6) mm thick, simple or with a few branches starting from near the base, with pointed apices or with narrow, regular cups non- or sparsely squamulose toward the base, continuously corticate, smooth to often wrinkled and longitudinally ridged, often somewhat glossy; transverse cracks and white medullary outgrowths often present at the base of podetia. Apothecia rare, brown, convex. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed, ovoid, developing along the margins of cups, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests and chemistry: two chemotypes (both C-, KC-, P+ red, UV-): 1) K- or K+ yellowish slowly turning brown (fumarprotocetraric acid complex and quaesitic acid). 2) K+ yellow (atranorin and fumarprotocetraric acid complex). Chemotype 1) is the most frequent.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather common
Subalpine belt: extremely common
Montane belt: absent
Dry submediterranean belt: absent
Humid submediterranean belt: absent
Padanian area: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (4561)
2002/12/18
tip of podetium with cup


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (17928)
2003/03/06


Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)


Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)


Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)


Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Cladonia_macroceras.htm
France, Col de l'Iseran - Savoie
5/10/2015


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Cladonia_macroceras.htm
France, Col de l'Iseran - Savoie
5/10/2015


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Cladonia_macroceras.htm
France, Col de l'Iseran - Savoie
5/10/2015


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Cladonia_macroceras.htm
France, Col de l'Iseran - Savoie
5/10/2015


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Cladonia_macroceras.htm
France, Col de l'Iseran - Savoie
5/10/2015


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=967&lang=en
France, Alpes Tarentaise 2000 m


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=967&lang=en
France, Alpes Tarentaise 2000 m


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=967&lang=en
France, Alpes Tarentaise 2000 m


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=967&lang=en
France, Alpes Tarentaise 2000 m


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=967&lang=en
France, Alpes Tarentaise 2000 m


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=967&lang=en
France, Alpes Tarentaise 2000 m


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C4/Cladonia_macroceras.htm
France, Col de l'Iseran - Savoie
5/10/2015


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Austria: Alps.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Puerto de Piedrafita (Asturias), a 1600 mt de altitud, entre briofitos, leg. & det. M. González, conf. A. R. Burgaz


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Großer Rachel


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, in alp. Cadubriae 1855
as C. gracilis


Triin Aimla; Owner: University of Tartu