Cladonia squamosa Hoffm. var. squamosa
Deutschl. Fl., 2: 125, 1796.
Synonyms: Cenomyce cucullata Delise; Cladonia squamosa var. muricella (Delise) Vain.; Cladonia squamosa var. phyllocoma Rabenh.; Cladonia squamosa var. polychonia Flörke
Description: Primary thallus squamulose, persistent or rarely disappearing, the squamules indented, 2-10 mm long, grey green above, white beneath, esorediate but sometimes granulose on the lower surface. Podetia extremely variable in shape and size, hollow inside, greenish grey to brown, not melanotic at base, 3-7(-14) cm tall, 0.5-2(-5) mm thick, simple or sparingly branched with open axils, usually expanded in upper parts to form gaping funnels which may proliferate radially, the surface often largely ecorticate exposing the stereome, more or less densely covered with microsquamules, squamules and granules, esorediate. Apothecia infrequent, brown, convex, located at the top of podetia or at the margin of funnels, often clustered. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed on podetial tips, rarely on basal squamules, with a red or colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV+ white. Chemistry: squamatic acid and variable amounts of barbatic acid (especially when fertile).
Growth form: Fruticose
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: rare
Montane belt: rare
Dry submediterranean belt: very rare
Humid submediterranean belt: very rare
Padanian area: extremely rare
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Current prediction (1981-2010)
Future prediction (2071-2100) SSP 1-2.6
Future prediction (2071-2100) SSP 5-8.5Predictive maps according to Francesconi et al. 2025
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15487)
2001/12/04

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15487)
2001/12/04
detail of cortex

Felix Schumm – CC BY-SA 4.0
[427], Germany, Baden-Württemberg, Kreis Göppingen, bei der Herrenmühle unterhalb Kloster Adelberg, Stubensandsteingebiet. Leg. et det. Schumm 30.03.1963, test. Klement 1967.

Felix Schumm – CC BY-SA 4.0
[427], Germany, Baden-Württemberg, Kreis Göppingen, bei der Herrenmühle unterhalb Kloster Adelberg, Stubensandsteingebiet. Leg. et det. Schumm 30.03.1963, test. Klement 1967.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[3152], Germany, Baden-Württemberg, Schwarzwald, nördlich von Beihingen westlich von Nagold, ca. 600 m, TK; 7417, leg. et det. F. Schumm 16.02.1988

Felix Schumm – CC BY-SA 4.0
[3152], Germany, Baden-Württemberg, Schwarzwald, nördlich von Beihingen westlich von Nagold, ca. 600 m, TK; 7417, leg. et det. F. Schumm 16.02.1988

Felix Schumm – CC BY-SA 4.0
[3152], Germany, Baden-Württemberg, Schwarzwald, nördlich von Beihingen westlich von Nagold, ca. 600 m, TK; 7417, leg. et det. F. Schumm 16.02.1988,
sq: squamatic acid

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Huelgoat

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Huelgoat

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Huelgoat
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Huelgoat
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Huelgoat
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.
Growth form: Fruticose
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: rare
Montane belt: rare
Dry submediterranean belt: very rare
Humid submediterranean belt: very rare
Padanian area: extremely rare
pH of the substrata:
1 | 2 | 3 | 4 | 5 |
Solar irradiation:
1 | 2 | 3 | 4 | 5 |
Aridity:
1 | 2 | 3 | 4 | 5 |
Eutrophication:
1 | 2 | 3 | 4 | 5 |
Poleotolerance:
0 | 1 | 2 | 3 |
Altitudinal distribution:
1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map



Predictive maps according to Francesconi et al. 2025

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15487)
2001/12/04

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15487)
2001/12/04
detail of cortex

Felix Schumm – CC BY-SA 4.0
[427], Germany, Baden-Württemberg, Kreis Göppingen, bei der Herrenmühle unterhalb Kloster Adelberg, Stubensandsteingebiet. Leg. et det. Schumm 30.03.1963, test. Klement 1967.

Felix Schumm – CC BY-SA 4.0
[427], Germany, Baden-Württemberg, Kreis Göppingen, bei der Herrenmühle unterhalb Kloster Adelberg, Stubensandsteingebiet. Leg. et det. Schumm 30.03.1963, test. Klement 1967.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[11332], Portugal, Madeira, Posto Florestal do Fanal zwischen Paúl da Serra und Ribeira da Janela; 32°48.545' N, 17°08.489' W, 1175 m; an alten bemoosten Laubbäumen. Leg. et det. Schumm 26.12.2003.

Felix Schumm – CC BY-SA 4.0
[3152], Germany, Baden-Württemberg, Schwarzwald, nördlich von Beihingen westlich von Nagold, ca. 600 m, TK; 7417, leg. et det. F. Schumm 16.02.1988

Felix Schumm – CC BY-SA 4.0
[3152], Germany, Baden-Württemberg, Schwarzwald, nördlich von Beihingen westlich von Nagold, ca. 600 m, TK; 7417, leg. et det. F. Schumm 16.02.1988

Felix Schumm – CC BY-SA 4.0
[3152], Germany, Baden-Württemberg, Schwarzwald, nördlich von Beihingen westlich von Nagold, ca. 600 m, TK; 7417, leg. et det. F. Schumm 16.02.1988,
sq: squamatic acid

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Huelgoat

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Huelgoat

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=411&lang=en
France, Forêt du Cranou

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Huelgoat
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Huelgoat
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Huelgoat
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1356&lang=en
France, Commana
form with brown interior of podetia
