Cladonia verticillata (Hoffm.) Schaer.
Lich. Helv. Spicil., 1, 1: 31, 1823. Basionym: Cladonia pyxidata (unranked) verticillata Hoffm. - Deutschl. Fl., 2: 122, 1796.
Synonyms: Cladonia cervicornis subsp. verticillata (Hoffm.) Ahti; Cladonia cervicornis var. verticillata (Hoffm.) Flot.; Cladonia verticillata var. evoluta (Th. Fr.) Stein
Description: Primary thallus squamulose, usually persistent and forming mats, the squamules very variable in size, 3-6 mm long, 2-5 mm wide, wedge-shaped or lobed, with indented margins, often ascending, upper surface grey-green to brownish in sun-forms, lower surface whitish, finely arachnoid. Podetia frequent, hollow inside, greenish grey to brown in sun-forms, 2-6(-8) cm tall, 0.5-3(-4) mm thick, flaring quickly into short, broad cups (up to 9 mm diam.), which tend to proliferate from the centre, often giving rise to up to 7 tiers of superimposed cups. Cortex more or less continuous to areolate, clearly maculate, the areoles smooth, subcontiguous. Apothecia infrequent, brown, convex, small. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, common at margins of apical cups, subglobose to ovoid, sessile, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K- or K+ yellowish slowly turning brown, C-, KC-, P+ red, UV-. Chemistry: fumarprotocetraric acid complex.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: extremely rare
Subalpine belt: rather rare
Montane belt: very rare
Dry submediterranean belt: extremely rare
Humid submediterranean belt: extremely rare
Padanian area: absent
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Roscanvel
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (3328)
2001/12/04
detail of proliferating cup
Felix Schumm – CC BY-SA 4.0
[15800], Australia, Western Australia, 75 km south of Mt. Chudalup, wetland with Agonis junipera and Eucalyptus marginata, growing on sunny rocks, 34.73380° S, 116.08760° E, 75 m. Leg. F. Schumm & E. Stocker-Wörgötter, 24.10.2009, det. T. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[15647], Australia, Western Australia, Perth Hills, east of Perth, 31.818° S, 116,456° E, 345 m, in dry Eucalyptus forest. Leg. F. Schumm & E. Stocker-Wörgötter, 21.10.2009, det. Schumm, conf. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[15800], Australia, Western Australia, 75 km south of Mt. Chudalup, wetland with Agonis junipera and Eucalyptus marginata, growing on sunny rocks, 34.73380° S, 116.08760° E, 75 m. Leg. F. Schumm & E. Stocker-Wörgötter, 24.10.2009, det. T. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[15800], Australia, Western Australia, 75 km south of Mt. Chudalup, wetland with Agonis junipera and Eucalyptus marginata, growing on sunny rocks, 34.73380° S, 116.08760° E, 75 m. Leg. F. Schumm & E. Stocker-Wörgötter, 24.10.2009, det. T. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti
Felix Schumm – CC BY-SA 4.0
[15647], Australia, Western Australia, Perth Hills, east of Perth, 31.818° S, 116,456° E, 345 m, in dry Eucalyptus forest. Leg. F. Schumm & E. Stocker-Wörgötter, 21.10.2009, det. Schumm, conf. Ahti 2011.,
p: protocetraric acid, f: fumarprotocetraric acid
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti,
fpr: fumarprotocetraric acid, pr: protocetraric acid
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Crozon
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Crozon
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Roscanvel
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Pointe de Treberon
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Commana
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Commana
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Commana
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.
Domenico Puntillo Cc by 4.9
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Roding
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Grafenwöhr
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Grafenwöhr
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Hochfels
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Hochfels
Triin Aimla; Owner: University of Tartu
originally as Cladonia verticillata
Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: extremely rare
Subalpine belt: rather rare
Montane belt: very rare
Dry submediterranean belt: extremely rare
Humid submediterranean belt: extremely rare
Padanian area: absent
pH of the substrata:
| 1 | 2 | 3 | 4 | 5 |
Solar irradiation:
| 1 | 2 | 3 | 4 | 5 |
Aridity:
| 1 | 2 | 3 | 4 | 5 |
Eutrophication:
| 1 | 2 | 3 | 4 | 5 |
Poleotolerance:
| 0 | 1 | 2 | 3 |
Altitudinal distribution:
| 1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Roscanvel
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (3328)
2001/12/04
detail of proliferating cup
Felix Schumm – CC BY-SA 4.0
[15800], Australia, Western Australia, 75 km south of Mt. Chudalup, wetland with Agonis junipera and Eucalyptus marginata, growing on sunny rocks, 34.73380° S, 116.08760° E, 75 m. Leg. F. Schumm & E. Stocker-Wörgötter, 24.10.2009, det. T. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[15647], Australia, Western Australia, Perth Hills, east of Perth, 31.818° S, 116,456° E, 345 m, in dry Eucalyptus forest. Leg. F. Schumm & E. Stocker-Wörgötter, 21.10.2009, det. Schumm, conf. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[15800], Australia, Western Australia, 75 km south of Mt. Chudalup, wetland with Agonis junipera and Eucalyptus marginata, growing on sunny rocks, 34.73380° S, 116.08760° E, 75 m. Leg. F. Schumm & E. Stocker-Wörgötter, 24.10.2009, det. T. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[15800], Australia, Western Australia, 75 km south of Mt. Chudalup, wetland with Agonis junipera and Eucalyptus marginata, growing on sunny rocks, 34.73380° S, 116.08760° E, 75 m. Leg. F. Schumm & E. Stocker-Wörgötter, 24.10.2009, det. T. Ahti 2011.
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti
Felix Schumm – CC BY-SA 4.0
[15647], Australia, Western Australia, Perth Hills, east of Perth, 31.818° S, 116,456° E, 345 m, in dry Eucalyptus forest. Leg. F. Schumm & E. Stocker-Wörgötter, 21.10.2009, det. Schumm, conf. Ahti 2011.,
p: protocetraric acid, f: fumarprotocetraric acid
Felix Schumm – CC BY-SA 4.0
[18963], Australia, New South Wales, Glenbog State Forest, ca. 7.6 km west of Bemboka on Snowy Mountains Highway, growing on soil in Eucalyptus woodland, 800 m. Leg. M.E. Hale (66366) 19.02.1984, det. T. Ahti,
fpr: fumarprotocetraric acid, pr: protocetraric acid
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C/Cladonia_cervicornis_vertic.htm
France, Paragan-Fenu - Corse
16/10/2015
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Crozon
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Crozon
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Kador
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Roscanvel
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Pointe de Treberon
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Commana
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Commana
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=300&lang=en
France, Commana
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Sweden: Värmland.
Domenico Puntillo Cc by 4.9
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Domenico Puntillo - CC BY 4.0
Italy, Calabria, Bosco di Mavigliano, Montalto Uffugo
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Roding
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Grafenwöhr
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Grafenwöhr
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Hochfels
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Hochfels
Triin Aimla; Owner: University of Tartu
originally as Cladonia verticillata
