Clavascidium lacinulatum (Ach.) M. Prieto

in Prieto & al., Am. J. Bot., 99: 28, 2012. Basionym: Endocarpon hepaticum var. lacinulatum Ach. - Lichenogr. Univ.: 299, 1810.
Synonyms: Catapyrenium lacinulatum (Ach.) Breuss; Dermatocarpon trapeziforme auct. non (J. König) Trevis.; Placidium lacinulatum (Ach.) Breuss
Description: Thallus squamulose, pale to dark brown, dull, non- or very slightly pruinose. Squamules , scattered to contiguous, rarely overlapping, 3-8 mm broad, 0.3-0.5 mm thick, at first rounded, then crenulate, flattened to weakly convex, smooth, with an undulate-crisped edge. Lower surface pale brown, sometimes darkening, attached by several, simple to weakly dichotomously branched, robust, 100-300 μm thick, several mm long, pale to brown rhizines, and by colourless, 3-5 μm thick rhizohyphae. Upper cortex paraplectenchymatous 40-80 µm thick; medulla of isodiametrical, thick-walled cells, well-delimited from lower cortex; lower cortex poorly developed, the cells not arranged in vertical rows. Perithecia frequent, laminal, immersed in the squamules, globose to broadly pyriform, up to 0.65 mm across, without involucrellum. Exciple colourless to pale brown throughout; hymenial gel hemiamyloid; hamathecium of periphyses measuring 25-35 x 2.5-3.5 µm, interascal filaments absent. Asci 8-spored, cylindrical, thin-walled, I-, without a tholus, 65-80 x 12-17 µm. Ascospores 1-celled, hyaline, ellipsoid, thin-walled, uniseriately arranged at least in young asci, 12-15(-17) x 5.5-7.5 μm. Pycnidia laminal, immersed, black, to 0.5 mm in diam., Dermatocarpon-type. Conidia oblong-ellipsoid, 3-4(-5) x 1.3-2 μm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: absent
Dry submediterranean belt: very rare
Humid submediterranean belt: very rare
Padanian area: absent

pH of the substrata:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

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Predictive map

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Glowacki
2003/04/28


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Glowacki
2003/04/28


Pier Luigi Nimis - CC BY-SA 4.0
TSB 41278


P.L. Nimis CC BY-SA 04
GZU Herb.Glowacki


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue, dunes


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Source: Zhang, T.; Zhang, X.; Yang, Q.; Wei, X. Hidden Species Diversity was Explored in Two Genera of Catapyrenioid Lichens (Verrucariaceae, Ascomycota) from the Deserts of China. J. Fungi 2022, 8, 729. https://doi.org/10.3390/jof8070729 - CC BY 4.0
China
The thallus habit and the anatomic structure of Clavascidium lacinulatum (HMAS-L 153962). (A). Squamulose thallus. (B). Transversal section of thallus. (C). Immersed perithecium. (D,E). Uniseriate ascospores in cylindrical asci. (F). Biseriate ascospores in clavate asci. (G). Conidia. Bars: (A) =2 mm, (B,C) =50 µm, (D,E) =10 µm, (F) =50 µm, and (G) =10 µm.